The ant genus Leptanilla: discovery of the worker-associated male of L. japonica, and a description of a new species from Taiwan (Hymenoptera: Formicidae: Leptanillinae)

The male and female of Leptanilla japonica Baroni Urbani are described. This is the first worker-associated male to be discovered in subfamily Leptanillinae. Leptanilla taiwanensis is described as new based on the worker and female from Taiwan, the first record of Leptanillinae from the island. An ecological note on the food of the ants is given. Implications of the male morphology and taxonomic positions of the other leptanilline genera based on males are discussed.